APS Member History

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1	Name:	Dr. Howard C. Berg
	Institution:	Harvard University
	Year Elected:	2002
	Class:	2. Biological Sciences
	Subdivision:	206. Physiology, Biophysics, and Pharmacology
	Residency:	Resident
	Living? :	Deceased
	Birth Date:	1934
	Death Date:	December 30, 2021

	Howard Berg received a Ph.D. at Harvard University in 1964, was a Junior Fellow of the Harvard Society of Fellows (1963-66), and remained at Harvard as an associate professor of biology and chairman of the Board of Tutors in Biochemical Sciences until 1970. He then moved to the University of Colorado, serving as professor of molecular, cellular and developmental biology and department chairman until 1979 when he became a professor of biology at the California Institute of Technology. In 1986 he returned to Harvard University and is currently professor of molecular and cellular biology and professor of physics, and a member of the Rowland Institute. Howard Berg was a chemist (B.S., Caltech); a medical student (two years at the Harvard Medical School); and then a graduate student in physics and Junior Fellow. As a physicist he worked with Norman Ramsey on the atomic hydrogen maser and with Ed Purcell on what is now called sedimentation field-flow fractionation. In 1968 he became interested in the motile behavior of bacteria. He has made many seminal contributions to understanding the biophysics of motility. Among other things, he and coworkers showed, via three-dimensional tracking, that E. coli executes a biased random walk and that bacterial flagella rotate: they do not wave or beat. This surprising conclusion has led Howard Berg to study the structure, genetics and physiology of the remarkable flagellar motor. Also, he has figured out how spirochetes swim, what bacterial flagella actually do when cells run and tumble and, with Ed Purcell, he developed the basic theory of the physics of chemoreception. His book Random Walks in Biology (1993), mostly about diffusion, has become a classic. His writings on life at low Reynolds numbers are great science and illustrate his intellectual reach: from pure physics to true biological understanding. His significant contributions to science reflect an approach to biological problems of a very perceptive biologist with the mind-set of a talented physicist. His inquisitiveness and productivity are models of scientific inquiry. A more recent book, E. coli in Motion (2004), reviews the field of bacterial chemotaxis. Dr. Berg is a member of the National Academy of Sciences and the American Academy of Arts & Sciences. In 2008 he received the Biophysical Society's annual award for Outstanding Investigator in Single Molecule Biology. He was elected a member of the American Philosophical Society in 2002.

2	Name:	Dr. Konrad E. Bloch
	Institution:	Harvard University
	Year Elected:	1966
	Class:	2. Biological Sciences
	Subdivision:	201. Molecular Biology and Biochemistry
	Residency:	Resident
	Living? :	Deceased
	Birth Date:	1912
	Death Date:	October 15, 2000

3	Name:	Dr. Barry R. Bloom
	Institution:	Harvard University
	Year Elected:	2004
	Class:	2. Biological Sciences
	Subdivision:	209. Neurobiology
	Residency:	Resident
	Living? :	Living
	Birth Date:	1937

	Barry Bloom's passion has been to relate the cutting edge of biomedical science to the needs of the 85% of the world's people living in resource-poor developing countries. His initial research analyzed the complex mechanisms of the immune response by developing in vitro models, enabling him to discover the first lymphokine or cytokine, non-antibody products of activated lymphocytes that regulate the functions of the immune system and mediate inflammation, tissue damage and resistance to microbial infection. After teaching the first course on immunology in India, he began research on leprosy. With collaborators, he created the first DNA library containing all the genes of the leprosy and the tubercle bacilli, thereby ultimately enabling the complete genomes of these major pathogens of humans to be sequenced. Those libraries and the first monoclonal antibodies produced against these pathogens were given to the World Health Organization (WHO) to distribute free of charge to scientists all over the world, helping to stimulate a global effort against these diseases. He has more recently explored the genetic basis of resistance of experimental animals against tuberculosis which integrates knowledge of the host and pathogen in understanding the disease. When there was a serious increase in tuberculosis in the U.S. in the early 1990s his group established, against conventional wisdom, that active transmission of infection, rather than reactivation of old infections, was an important component of the epidemic. Such transmission required implementation of stringent public health measures. He has worked in an official capacity for the WHO for the past 37 years and has advised the National Institutes of Health, the National Academy of Sciences and the White House on scientific issues and on international health policies. Dr. Bloom is currently Dean of the Faculty of the Harvard School of Public Health.

4	Name:	Dr. Lawrence Bogorad
	Institution:	Harvard University
	Year Elected:	1985
	Class:	2. Biological Sciences
	Subdivision:	202. Cellular and Developmental Biology
	Residency:	Resident
	Living? :	Deceased
	Birth Date:	1921
	Death Date:	December 28, 2003

5	Name:	Dr. Emery N. Brown
	Institution:	Massachusetts Institute of Technology; Harvard Medical School; Massachusetts General Hospital
	Year Elected:	2023
	Class:	2. Biological Sciences
	Subdivision:	204. Medicine, Surgery, Pathology and Immunology
	Residency:	Resident
	Living? :	Living
	Birth Date:	1956

	Emery N. Brown, M.D., Ph.D. was born and raised in Ocala, Florida. He attended Fessenden Elementary School, Osceola Junior High School and North Marion High School before graduating from the Phillips Exeter Academy in Exeter, New Hampshire. He spent the second semester of his senior year in Barcelona, Spain studying Spanish with the School Year Abroad Program. Brown received his B.A. in Applied Mathematics (magna cum laude) from Harvard College. Before entering the Harvard M.D. Ph.D. Program, he spent a year studying mathematics as a Rotary Fellow at the Fourier Institute in Grenoble, France. Brown went on to earn his M.A. and Ph.D. in statistics from Harvard University and his M.D. (magna cum laude) from Harvard Medical School. He completed his internship in internal medicine at the Brigham and Women’s Hospital and his anesthesiology residency at MGH. After completing his residency in 1992, Brown joined the faculty at the MGH Department of Anesthesia and Harvard Medical School. In 2005, he also joined the faculty at MIT in the Department of Brain and Cognitive Sciences and the MIT and Harvard Health Sciences and Training Program. Brown is presently the Edward Hood Taplin Profess of Medical Engineering and Computational Neuroscience at MIT; the Warren M. Zapol Professor of Anaesthesia at Harvard Medical School; and an anesthesiologist at MGH. He is an anesthesiologist-statistician whose research is defining the neuroscience of how anesthetics produce the states of general anesthesia. He also develops statistical methods for neuroscience data analysis. In 2013 to 2014, Brown served on President Obama’s Brain Initiative Working Group. Currently, he serves on the Board of Trustees for the Simons Foundation and the Guggenheim Foundation. Brown is a fellow of the American Institute for Medical and Biological Engineering, the IEEE, the American Statistical Association, the Institute of Mathematical Statistics, the American Academy of Arts and Sciences and the National Academy of Inventors. Brown is also a member of the National Academy of Medicine, the National Academy of Sciences, and the National Academy of Engineering. Brown is the recipient of an NIH Director’s Pioneer Award, a Guggenheim Fellowship in Applied Mathematics, the American Society of Anesthesiologists Excellence in Research Award, the Dickson Prize in Science, the Swartz Prize for Theoretical and Computational Neuroscience, the Pierre Galletti Award, the Gruber Prize in Neuroscience and Doctorates of Science Honoris Causas from the University of Southern California and SUNY Downstate.

6	Name:	Dr. William B. Castle
	Institution:	Harvard University
	Year Elected:	1939
	Class:	2. Biological Sciences
	Subdivision:	204. Medicine, Surgery, Pathology and Immunology
	Residency:	Resident
	Living? :	Deceased
	Birth Date:	1897
	Death Date:	8/9/90

7	Name:	Dr. Paul Mead Doty
	Institution:	Harvard University
	Year Elected:	1970
	Class:	2. Biological Sciences
	Subdivision:	201. Molecular Biology and Biochemistry
	Residency:	Resident
	Living? :	Deceased
	Birth Date:	1920
	Death Date:	December 5, 2011

	Paul M. Doty was professor of public policy and Mallinckrodt Professor of Biochemistry Emeritus at Harvard University's John F. Kennedy School of Government. He was also the founder and Director Emeritus of the Center for Science and International Affairs and an emeritus member of the BCSIA Board of Directors. During his 42 years on the Harvard University faculty, Dr. Doty embraced two careers: one in biochemistry, where he founded the Department of Biochemistry and Molecular Biology, and the other in science policy and international security studies, where he founded the Center for Science and International Affairs in 1974. From 1960-64 he was a member of the President's Science Advisory Committee and from 1961-67 he also served as a consultant to the United States Arms Control and Disarmament Agency. The recipient of awards including the American Chemical Society Award in Pure Chemistry, Dr. Doty was the author of numerous articles in scientific journals and is a member of the National Academy of Sciences. He was elected to membership in the American Philosophical Society in 1970. Paul Doty died on December 5, 2011, at age 91 at his home in Cambridge, Massachusetts.

8	Name:	Dr. John E. Dowling
	Institution:	Harvard University
	Year Elected:	1992
	Class:	2. Biological Sciences
	Subdivision:	208. Plant Sciences
	Residency:	Resident
	Living? :	Living
	Birth Date:	1935

	John E. Dowling received his A.B. and Ph.D. from Harvard University. He taught in the Biology Department at Harvard from 1961 to 1964, first as an Instructor, then as Assistant Professor. In 1964 he moved to Johns Hopkins University, where he held an appointment as Associate Professor of Ophthalmology and Biophysics. He returned to Harvard as Professor of Biology in 1971, was the Maria Moors Cabot Professor of Natural Sciences from 1971-2001, Harvard College Professor from 1999-2004 and is presently the Gordon and Llura Gund Professor of Neurosciences. He was Chairman of the Biology Department at Harvard from 1975 to 1978 and served as Associate Dean of the Faculty of Arts and Sciences from 1980 to 1984. He was Master of Leverett House at Harvard from 1981-1998 and served as President of the Corporation of The Marine Biological Laboratory in Woods Hole from 1998-2008. He is a Fellow of the American Academy of Arts and Sciences, a member of the National Academy of Sciences and a member of the American Philosophical Society. He received the Friedenwald Medal from the Association of Research in Ophthalmology and Vision in 1970, the Annual Award of the New England Ophthalmological Society in 1979, the Retinal Research Foundation Award for Retinal Research in 1981, an Alcon Vision Research Recognition Award in 1986, a National Eye Institute’s MERIT award in 1987, the Von Sallman Prize in 1992, The Helen Keller Prize for Vision Research in 2000, the Llura Ligget Gund Award for Lifetime Achievement and Recognition of Contribution to the Foundation Fighting Blindness in 2001 and the Paul Kayser International Award in Retina Research in 2008.. He was granted an honorary M.D. degree by the University of Lund (Sweden) in 1982. His research interests have focused on the vertebrate retina as a model piece of the brain. He and his collaborators have long been interested in the functional organization of the retina, studying its synaptic organization, the electrical responses of the retinal neurons, and the mechanisms underlying neurotransmission and neuromodulation in the retina. He became interested in zebrafish as a system in which one could explore the development and genetics of the vertebrate retina about 15 years ago. Part of his research team has focused on retinal development in zebrafish and the role of retinoic acid in early eye and photoreceptor development. A second group has developed behavioral tests to isolate mutations, both recessive and dominant, specific to the visual system.

9	Name:	Dr. Andrew H. Knoll
	Institution:	Harvard University
	Year Elected:	1997
	Class:	2. Biological Sciences
	Subdivision:	203. Evolution & Ecology, Systematics, Population Genetics, Paleontology, and Physical Anthropology
	Residency:	Resident
	Living? :	Living
	Birth Date:	1951

	Andrew Knoll is Fisher Professor of Natural History and Professor of Earth and Planetary Sciences at Harvard University and Curator of the Paleobotanical Collections in the Harvard University Herbaria. A world leader in paleobiology, he has served on the Harvard faculty since 1982. Distinguished by his pioneering investigations of global conditions in the early evolution of life, Dr. Knoll has used biological, geological and chemical information to interpret the evolution of unicellular and multicellular organisms, from the most primitive life forms to the origin of metazoa and higher plants. He has used the same approach to explain mass extinctions in the Permian. Dr. Knoll's many honors include the Walcott Medal (1987) and the Mary Clark Thompson Medal (2012) of the National Academy of Sciences, the Phil Beta Kappa Book Award in Science (2003), the Moore Medal of the Society for Sedimentary Geology (2005), the Paleontological Society Medal (2005), the Wollaston Medal of the Geological Society of London (2007), and membership in the National Academy of Sciences and the American Academy of Arts & Sciences. He received his Ph.D. from Harvard in 1977.

10	Name:	Dr. Catherine Dulac
	Institution:	Harvard University; Howard Hughes Medical Institute
	Year Elected:	2019
	Class:	2. Biological Sciences
	Subdivision:	208. Plant Sciences
	Residency:	Resident
	Living? :	Living
	Birth Date:	1963

	Catherine Dulac is a Howard Hughes Medical Institute Investigator, Higgins Professor of Molecular and Cellular Biology and Lee and Ezpeleta Professor of Arts and Sciences at Harvard University. Her work explores the identity and function of neural circuits underlying instinctive social behaviors in mice, and the role of genomic imprinting in the adult and developing brain. She grew up in Montpellier, France, graduated from the Ecole Normale Supérieure, Paris, and received her PhD from the University of Paris VI. She was a postdoctoral fellow at Columbia University and joined the faculty of Harvard as a junior faculty in 1996, before becoming full professor in 2001, and Chair of Harvard's Department of Molecular and Cellular Biology from 2007 until 2013. She is a member of the US National Academy of Sciences, and of the French Academy of Sciences, and a fellow of the American Academy of Arts and Sciences and the American Association for the Advancement of Science. She is a recipient of multiple awards including the Richard Lounsbery Award, the National Academy’s Pradel Research Award, the Edward M. Scolnick Prize in Neuroscience, the Karl Spencer Lashley Award from the American Philosophical Society, and the 2021 Breakthrough Prize for Life Sciences.

11	Name:	Dr. John T. Edsall
	Institution:	Harvard University
	Year Elected:	1955
	Class:	2. Biological Sciences
	Subdivision:	201. Molecular Biology and Biochemistry
	Residency:	Resident
	Living? :	Deceased
	Birth Date:	1902
	Death Date:	June 12, 2002

12	Name:	Dr. Scott Edwards
	Institution:	Harvard University
	Year Elected:	2020
	Class:	2. Biological Sciences
	Subdivision:	203. Evolution & Ecology, Systematics, Population Genetics, Paleontology, and Physical Anthropology
	Residency:	resident
	Living? :	Living
	Birth Date:	1963

	Scott Edwards is currently Alexander Agassiz Professor, Curator of Birds at the Museum of Comparative Zoology, and Professor of Organismic and Evolutionary Biology at Harvard University. He earned his Ph.D. from the University of California, Berkeley in 1992, after which he spent a decade at the University of Washington. Scott Edwards pioneered the application of genomics and population genetic approaches to natural populations of birds. He creatively integrates theory with field, museum, and laboratory research. Edwards has led his field of ornithology into the age of genomics by examining genomic evolution in a setting that applies population genetics, systematics, and natural history to diverse questions. His work has broad generality beyond the target taxa, and he has contributed importantly to both theory and methods. His lab applies cutting edge population genetics to studies on diverse problems in genomics and evolutionary biology. With Beerli he modernized phylogeography. With Liu and Pearl he developed methods that promise to revolutionize phylogenetic analyses of large molecular datasets. He has made extensive contributions to understanding behavioral evolution, speciation, and biogeography of Australian birds. Edwards is a versatile and prolific scientist who has proven to be an effective communicator and a wonderful role model. In 2015 he won the Elliot Coues Award of the American Ornithologists Union. He is a member of the Society of Systematic Biology (president, 2009), the American Academy of Arts & Sciences (2009), the American Genetic Association (president, 2011), and the National Academy of Sciences (2015). Scott Edwards was elected a member of the American Philosophical Society in 2020.

13	Name:	Dr. Judah Folkman
	Institution:	Harvard Medical School & Children's Hospital
	Year Elected:	1999
	Class:	2. Biological Sciences
	Subdivision:	204. Medicine, Surgery, Pathology and Immunology
	Residency:	Resident
	Living? :	Deceased
	Birth Date:	1933
	Death Date:	January 14, 2008

14	Name:	Dr. Laurie H. Glimcher
	Institution:	Dana Farber Cancer Institute; Harvard Medical School
	Year Elected:	2019
	Class:	2. Biological Sciences
	Subdivision:	209. Neurobiology
	Residency:	Resident
	Living? :	Living
	Birth Date:	1951

	Laurie H. Glimcher is President and Chief Executive Officer of the Dana-Farber Cancer Institute and Richard and Susan Smith Professor of Medicine at Harvard Medical School. She earned her M.D. from Harvard Medical School in 1976, where she spent most of her career, including as Irene Heinz Given Professor of Immunology. Laurie Glimcher has elucidated the molecular pathways that regulate the development and activation of cells in the immune system - pathways critical for both the development of protective immunity and for the pathophysiologic immune responses underlying autoimmune, infectious, allergic, and malignant diseases. She discovered the first Th1-specific transcription factor, T-bet, and demonstrated that it is the master-regulator of Type 1 immunity in cells of both the adaptive and innate immune system. She also discovered XBP1, the first transcription factor required for both plasma cell differentiation and the Endoplasmic Reticulum (ER) Stress Response. She then demonstrated a link between ER stress and proinflammatory/autoimmune diseases. Most recently she discovered that XBP1 is key in the maintenance of cancer stem cells in triple negative breast cancer. Further, IRE1/XBP1 also controls anti-tumor immunity by disrupting dendritic cell homeostasis. Hence reducing IRE1/XBP1 activity should simultaneously inhibit tumor cell growth and activate type 1 anti-tumor immunity. She is a member of the American Academy of Arts & Sciences (1996), the National Academy of Sciences (2002), and the American Association of Immunologists, (president, 2003-04). Laurie Glimcher was elected a member of the American Philosophical Society in 2019.

15	Name:	Dr. Stephen Coplan Harrison
	Institution:	Harvard University & Howard Hughes Medical Institute
	Year Elected:	1997
	Class:	2. Biological Sciences
	Subdivision:	206. Physiology, Biophysics, and Pharmacology
	Residency:	Resident
	Living? :	Living
	Birth Date:	1943

	Stephen Harrison is a world leader in understanding virus structure and in probing the relationship between structure and function of complex protein assemblies. He has devised innovative methodological enhancements of X-ray crystallography to determine the detailed molecular structures of viruses, cell-surface receptors, and DNA-protein complexes. His pioneering studies of small plant viruses at atomic dimensions revealed the basic molecular design of a large class of RNA viruses of plants, insects, and vertebrates. This work, and his subsequent studies of many other viruses, allowed Dr. Harrison to formulate principles that govern viral structure, assembly, stability, cellular attachment, and fusion. This information is fundamental to understanding viral disease and to the design of antiviral drugs and vaccines. Similarly, Dr. Harrison's X-ray crystallographic studies of the structures of DNA-protein complexes revealed important molecular mechanisms in the control of gene activity. Dr. Harrison earned his Ph.D. from Harvard University in 1967. He joined the Harvard faculty in 1971 and is currently director of the Center for Molecular and Cellular Dynamics at Harvard Medical School and head of the Laboratory of Molecular Medicine at Children's Hospital, Boston. He won the Welch Award in Chemistry in 2015 and the Rosenstiel Award for Basic Medical Research in 2018.

16	Name:	Dr. Hopi E. Hoekstra
	Institution:	Harvard University; Howard Hughes Medical Institute
	Year Elected:	2018
	Class:	2. Biological Sciences
	Subdivision:	203. Evolution & Ecology, Systematics, Population Genetics, Paleontology, and Physical Anthropology
	Residency:	Resident
	Living? :	Living
	Birth Date:	1972

	Hopi E. Hoekstra is the Alexander Agassiz Professor of Zoology in the Departments of Organismic & Evolutionary Biology and the Molecular & Cellular Biology at Harvard University. She is the Curator of Mammals in the Museum of Comparative Zoology, an Institute Member at the Broad Institute and an Investigator at the Howard Hughes Medical Institute. Her research focuses on understanding the evolution and genetics of morphological and behavioral traits that affect fitness of individuals in the wild. Using deer mice as a model system, she first dissected the molecular, genetic and developmental basis of camouflaging coloration to understand the mechanisms driving adaptation. Later, she focused on unraveling the genetic and neurobiological underpinnings of complex natural behaviors. She received her B.A. from UC Berkeley and her Ph.D. from the University of Washington. She has received Young Investigator awards from the American Society of Naturalists and the Arnold and Mabel Beckman Foundation, and most recently, the Lounsbery Medal from the National Academy of Sciences (2015). She gave the 2013 Commencement speech at UC Berkeley’s Integrative Biology Department and has been profiled in The New York Times. In 2016, she was elected into the National Academy of Sciences and in 2017, the American Academy of Arts & Sciences. She also teaches in Harvard’s introductory Life Science course Genetics, Genomics and Evolution to approximately 500 freshmen each year, and has been awarded the Fannie Cox Prize and a Harvard College Professorship for teaching excellence.

17	Name:	Dr. David H. Hubel
	Institution:	Harvard University
	Year Elected:	1982
	Class:	2. Biological Sciences
	Subdivision:	209. Neurobiology
	Residency:	Resident
	Living? :	Deceased
	Birth Date:	1926
	Death Date:	September 22, 2013

	David Hubel received the 1981 Nobel Prize in Physiology or Medicine, along with Torsten Wiesel, for his pioneering work on the functioning of the visual system of mammals. His studies have shown how the visual cortex develops physiologically and how it records what the eye sees. This work has led to new understanding and treatment of childhood eye afflictions and to studies of cortical plasticity. Born in Ontario, Canada in 1926, Dr. Hubel received his M.D. from McGill University in 1951. He worked at Johns Hopkins Hospital from 1952-59 and at Walter Reed Hospital, where he began comparing the activity of sensory cells in waking and sleeping animals. Dr. Hubel had been a member of the Harvard Medical School faculty since 1959 and was Research Professor of Neurobiology at Harvard University at the time of his death. David Hubel died September 22, 2013, at age 87, in Lincoln, Massachusetts.

18	Name:	Dr. Eugene Patrick Kennedy
	Institution:	Harvard University
	Year Elected:	1993
	Class:	2. Biological Sciences
	Subdivision:	201. Molecular Biology and Biochemistry
	Residency:	Resident
	Living? :	Deceased
	Birth Date:	1919
	Death Date:	September 22, 2011

	Eugene Patrick Kennedy was born in Chicago in 1919. He enrolled at De Paul University in 1937 as a chemistry major and then went to the University of Chicago in 1941 for graduate training in organic chemistry. To pay his tuition, Dr. Kennedy also got a job in the chemical research department of Armour and Company, one of the large meat packers in Chicago. As part of the war effort, his job at Armour was to assist in the large scale fractionation of bovine blood to obtain pure bovine serum albumin. It was believed that the bovine serum albumin might be useful for treating shock in soldiers on the battlefield. However, by the end of 1942, hope had faded that bovine serum albumin would be an effective treatment, and the Red Cross started to collect blood from volunteers instead. Armour opened a new facility in Fort Worth, Texas for the fractionation of human blood from donors, and Kennedy was sent to Fort Worth to assist in this effort. He remained in Texas until 1945, when the war was nearing its end and large amounts of human plasma proteins had been stockpiled. Returning to the University of Chicago, Dr. Kennedy immediately transferred from the Department of Chemistry to the Department of Biochemistry. His experience on the plasma project had led to a new appreciation of biochemistry. After finishing graduate school, Dr. Kennedy went to the University of California, Berkeley, to work with Horace A. Barker, who had just discovered that soluble extracts of Clostridium kluyveri cells could produce short-chain fatty acids from ethyl alcohol. Although the initial discovery had already been made, there was much to be learned about these extracts, and Dr. Kennedy aided in this effort. In 1950, he joined Fritz Lipmann at Harvard Medical School. He then returned to the University of Chicago in 1951, after being given a joint appointment in the Department of Biochemistry and the newly organized Ben May Laboratory for Cancer Research. In 1959, he was invited to become a Hamilton Kuhn Professor and head of the Department of Biological Chemistry at the Harvard Medical School. Over the course of his career, Dr. Kennedy made major contributions to the biosynthesis of phospholipids, the basic component of all membranes, and to our understanding of membrane function. He discovered the first step of phospholipid synthesis, the reaction of cytidine triphosphate and phosphorylcholine to form cytidine diphosphocholine, as well as the enzyme which catalyzes the reaction. It was Dr. Kennedy who found that a protein, permease, was responsible for the transport of sugars through the bacterial membrane. His research consistently elucidated the structure, localization and biosynthesis of oligosaccharides derived from membranes. Dr. Kennedy's interests also led him to investigate membrane biogenesis and function in bacteria, the translocation of membrane phospholipids, and periplasmic glucans and cell signaling in bacteria. He was the recipient of many honors including the Gairdner Foundation Award and the American Chemical Society's Paul Lewis Award. He was a member of the American Academy of Arts & Sciences and the National Academy of Sciences. Eugene Patrick Kennedy died on September 22, 2011, at the age of 92 in Cambridge, Massachusetts. Dr. Kennedy was at Harvard as the Hamilton Kuhn Professor of Biological Chemistry and Molecular Pharmacology, Emeritus.

19	Name:	Dr. Seymour S. Kety
	Institution:	Harvard University
	Year Elected:	1975
	Class:	2. Biological Sciences
	Subdivision:	209. Neurobiology
	Residency:	Resident
	Living? :	Deceased
	Birth Date:	1915
	Death Date:	May 25, 2000

20	Name:	Dr. Marc Kirschner
	Institution:	Harvard University
	Year Elected:	2021
	Class:	2. Biological Sciences
	Subdivision:	202. Cellular and Developmental Biology
	Residency:	Resident
	Living? :	Living
	Birth Date:	1945

	Marc Kirschner is the John Franklin Enders University Professor at Harvard University. He earned his Ph.D. from the University of California, Berkeley in 1971. Prior to arriving at Harvard, he taught at Princeton University from 1972 to 1978 and the University of California, San Francisco from 1978 to 1993. In 1993, he moved to Harvard Medical School (HMS), where he served as the Chair of the new Department of Cell Biology for a decade. He became the Founding Chair of the HMS Department of Systems Biology in 2003. Kirschner pioneered at least three fundamental and general concepts that help explain how biology organizes information spatially and temporally. He is a biochemist by training, but has always had a strong interest in using mathematics and physical principles to understand biology at a deeper level. In his game-changing research on the cytoskeleton, Kirschner discovered that microtubules explore space randomly and selectively reinforce productive connections, a concept at the crux of connectivity in the brain, angiogenesis, and many other processes. In his work on the cell cycle, he identified an autonomous oscillation that entrains the order of downstream events. The circadian clock and the vertebrate somite segmentation clock use similar principles. In frog embryo development, he found that a locally produced factor, FGF, provides instructions that induce a region of tissue to adopt a new fate; this discovery informed much of our understanding of developmental patterning. These seminal discoveries, and the technologies he developed to enable them, have been profoundly influential throughout biology and establish him as one of the great experimental biologists of all time. Kirschner has also been an advocate for federal biomedical research funding and served as first chair of the Joint Steering Committee for Public Policy, a coalition of scientific societies he helped create in 1993 to educate the U.S. Congress on biomedical research and lobby for public funding of it. Kirschner helped launch the monthly, peer-reviewed journal PLoS Biology in October 2003 as a member of the editorial board and senior author of a paper in the inaugural issue. He received the Richard Lounsbery Award in 1991, the Gairdner Foundation International Award in 2001, the American Society for Cell Biology's E.B. Wilson Medal in 2003, Carnegie Mellon University's Dickson Prize for Science in 2004, and Technion's Harvey Prize in 2015, the American Society for Cell Biology's Public Service Award in 1996, the William C. Rose Award, American Society for Biochemistry and Molecular Biology in 2001, the Rabbi Shai Shacknai Memorial Prize in Immunology and Cancer Research in 2003, and Carnegie Mellon University's Dickson Prize for Science in 2004. He was President of the American Society for Cell Biology from 1990 to 199. He has been a member of the National Academy of Sciences since 1989, the American Academy of Arts & Sciences since 1989, the Royal Society of London since 1999, and the Academia Europaea since 1999. Kirschner was elected a member of the American Philosophical Society in 2021.

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